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The player takes control of various members of Task Force Vanguard, a team formed by the Special Operations Executive to uncover Phoenix, a secret Nazi project. The game portrays the rise of special forces, with the first tasked to uncover a secret Nazi project and thwart it. The flashbacks portray various major battles such as the Battles of Stalingrad and Midway through the eyes of the main protagonists. The story of Vanguard is told throughout various points in time in a non-chronological order. Vanguard's single-player campaign takes place in 1945 near the end of World War II's European theater, with various flashbacks ranging from 1941 to 1944. Classic round-based maps were announced for Vanguard in April 2022, with the first map, a remake of the Call of Duty: World at War map 'Shi no Numa', slated for release in Season 4. An additional hub map, 'Terra Maledicta', was also added to the game in February 2022 as part of the Season 2 content update. Post-launch updates for Zombies feature additional objectives for Der Anfang, including a main quest for story progression, as well as new Covenants and field upgrades. A new gameplay feature, the Altar of Covenants, allows players to purchase randomized buffs every round, allowing a variety in combat builds. Gameplay elements from Cold War return in Vanguard, such as Essence and Salvage currencies, and Dark Aether-powered field upgrades, in addition to classic Zombies gameplay items like perks and the Pack-a-Punch machine. At launch, the game features a new map titled 'Der Anfang', which combines aspects of round-based survival gameplay with the objective-based gameplay of Outbreak and Onslaught, both new gamemodes introduced in Cold War.

The mode is considered an expansion of the Dark Aether story, and acts as a prologue to Black Ops Cold War's story. The cooperative Zombies mode returns in Vanguard, developed by Treyarch in collaboration with Sledgehammer Games. Like prior Call of Duty installments since 2019, Vanguard's multiplayer is also fully cross-platform play compatible, with support for cross-generation gaming regarding eighth and ninth generation home video game consoles. Īs with Cold War, Vanguard is integrated with Warzone, allowing players to progress and use weapons, operators and other cosmetics items across both titles, in addition to the existing Modern Warfare and Cold War items in Warzone. As an addition to the returning Gunsmith system, Vanguard 's multiplayer features a 'Caliber system', which brings greater destructibility and reactive environments to maps through different ammunition types. This allows the player to have more control over the selection of multiplayer game modes through three pre-defined pacing filters favoring different playstyles.

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A new feature introduced to the Call of Duty series through Vanguard's multiplayer is 'Combat Pacing'. Season 2 of Vanguard introduces 'Arms Race', a large-scale game mode where two teams of 12 players attempt to capture five bases, or destroy them players can also earn cash from kills and complete objectives to buy custom loadouts and killstreaks. Champion Hill tasks players with surviving as long as possible in a squad-based deathmatch, round-robin tournament. Vanguard's multiplayer mode introduces a new game mode to the series, titled 'Champion Hill', a successor to the mode 'Gunfight', a 2v2 arena mode previously featured in Call of Duty: Modern Warfare and Call of Duty: Black Ops Cold War. New gameplay features allow the player the ability to use a more advanced tactical approach in combat such as blind firing from behind cover, breaking through destructible environmental elements or creating new paths to complete objectives by climbing walls. Vanguard's campaign features similar gameplay mechanics previously introduced in Modern Warfare, such as the player being able to mount wielded weapons to flat surfaces, interact with doors and execute takedowns.

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